404. That’s an error.

The requested URL /bookmarks/mark?op=add&bkmk=http%3A//jcs.biologists.org/content/early/2019/03/11/jcs.224378&title=Multiple%20feedback%20mechanisms%20fine-tune%20Rho%20signaling%20to%20regulate%20morphogenetic%20outcomes was not found on this server. That’s all we know.