THE RAFFLES BULLETIN OF ZOOLOGY 2009 THE RAFFLES BULLETIN OF ZOOLOGY 2009 Supplement No. 20: 291–307 Date of Publication: 31 Jul.2009 © National University of Singapore THE RAFFLES BULLETIN, 1928–2009: EIGHT DECADES OF BRACHYURAN CRAB RESEARCH (CRUSTACEA: DECAPODA) Martyn E. Y. Low and S. H. Tan Raffles Museum of Biodiversity Research, Department of Biological Sciences, National University of Singapore, Block S6, Science Drive 2, Singapore 117546, Republic of Singapore Email: martyn@nus.edu.sg (Corresponding author) Peter K. L. Ng Tropical Marine Science Institute, National University of Singapore, 14 Kent Ridge Road, Singapore 119223 ABSTRACT. – The Raffles Bulletin of Zoology, and its predecessors (the Bulletin of the Raffles Museum and the Bulletin of the National Museum), have published a higher than average number of papers relating to brachyuran crabs. Although most deal with taxonomy and systematics, there were several papers dealing with ecology, larval development, ecology, on historical collections of specimens and even a molecular population genetics analysis. In this paper, we have compiled a list of these papers over eighty years. This will provide a standalone reference relevant to brachyuran taxonomy. Taxonomic and systematic descriptions and decisions are the main focus of this article but a complete list of papers is given. A total of 300 names have been published in the Bulletin (and its predecessors). Of these, three are replacement names, 17 have been synonymised and the remainder are valid. In addition to this, there have been changes to the dates of publications for several volumes of the Bulletin, pushing back the year of publication of 12 volumes by a year and and bringing forward one of them by a year. KEY WORDS. – Brachyura, Raffles Bulletin of Zoology, Bulletin of the Raffles Museum, Bulletin of the National Museum. INTRODUCTION Started in 1928, first as the Bulletin of the Raffles Museum, and later the Bulletin of National Museum, to publish studies on the collections of the Raffles Museum, the Raffles Bulletin of Zoology has seen as many changes as the country and its parent institution. Originally an organ of the Raffles Museum, Southeast Asia’s second oldest natural history museum; the Bulletin was subsequently ‘nationalised” when the country became independent in 1960 and the institution became the National Museum of Singapore. When the National Museum was “reorganised” in 1970 and the natural history collections moved out to the then University of Singapore (present National University of Singapore), the zoological component became the Zoological Reference Collection (ZRC). In 1988, the Raffles Bulletin was resurrected by entomologist Dennis Murphy, and it assumed its original tasks once again. With the “resurrection” of the Museum as the Raffles Museum of Biodiversity research in 1998, the Raffles Bulletin of Zoology became the organ of the new institute (see Ng et al. 1994)! The Museum’s association with decapod crustaceans and in particular, brachyuran crabs has been a long one. The first study on Brachyura was published just four years after the Bulletin got of to a start—with N. Smedley’s study on Parathelphusa sexpunctatum in the Malay Peninsula taking the honour. The first descriptions of new species were those by Swiss biologist Jean Roux (1935), the American Melbourne Ward (1935), and the Museum’s resident zoologist, Michael Tweedie (1935). Anchored by Michael Tweedie from the late 1930s, the brachyuran research activity was an active one as he had a deep passion for crabs (Ng & Yang, 1989; Ng, 1995b). He not only studied crabs himself but also sent material to specialists all over the world for study, some of which were published in the Bulletin. In the 1960s, then French UNESCO Taxonomy Expert, Raoul Serène, was based in Singapore for many years, adding substantially not only to its large collections, but also training students and sending material abroad. As one of the most prolific and proficient carcinologists (decapod crustacean expert) of the 20th century (Crosnier, 1982), his impact of the brachyuran collections and research was very substantial. At about the same time, English carcinologist and limnologist Desmond Johnson became staff at the then University of Singapore, 291 Low et al.: Brachyuran crab research in the Raffles Bulletin and in the next decade, added substantially to the material when his collections were integrated with the Museum’s in the late 1970s. This heritage has not been lost on the present generation of researchers. The extensive collections and literature of the early carcinologists like Tweedie, Serène and Johnson have set the stage for recent workers to work from where they left off. Large numbers of university students have also use these resources to complete their dissertations, and a good number of the papers in the Bulletin are from them. In addition, because of the research activity and resources available, numerous carcinologists from all over the world have used the museum's material for studies, and a good number have also visited the collection and collaborated with researchers and students in the university. In turn, carcinologists from the Museum have participated in numerous regional expeditions (in turn collecting more material), as well as worked on the crab fauna from dozens of countries all over the Indo-West Pacific. Many of the papers published in the Bulletin are the results of these expeditions, collaborations and loans. And the impact the Bulletin has had on carcinology has also been substantial. Since those early days, the Bulletin has published 305 species names (Table 1). Three are replacement names (two valid and one synonymised) and 17 have been synonymised. The 285 valid species first described in the Bulletin represent an average of 3.48 new species described every year over the past 82 years. In addition, 99 genera have been published during this same period (Table 2). One was a replacement name, and of the remaining 98, only two have been synonymised. The remaining 96 genera first described in the Bulletin are still recognised (Ng et al., 2008). Above the generic level, four subfamilies (Antrocarcininae Ng & D. G. B. Chia, 1994, Gaeticinae Davie & N. K. Ng, 2007a, Kraussiinae Ng, 1993, Thalassograpsinae, Davie & N. K. Ng, 2007a) and the family Xenograpsidae N. K. Ng, Davie, Schubart & Ng, 2007, have been was described in the Bulletin (Table 3). RESOURCES ON- AND OFFLINE In this short analysis, we seek to bring together all the literature relevant to brachyuran crabs published in the Bulletin in one place, as it were, to make it easier for workers to find this veritable mine of information. The other half of this effort is virtual. The Bulletin as with other journals of a certain age, have had to make the transition from paper to electronic formats and media. Many of the older versions of these digitised resources have been of a rather poor standard—a first pass done when getting the Bulletin online in its entirety took precedence. Efforts are now under way to rectify this and to replace the older PDFs with properly scanned text, figures and especially plates (the bread and butter of taxonomy). The scans will also be sufficiently high to allow for optical character recognition (OCR) and text search functions via the Adobe™ PDF application/plugin. With the publication of Ng, Guinot & Davie’s Supplement 17 of this Bulletin (Systema Brachyurorum I) in 2008, a complete placement of all extant brachyura was made available. We would be among the first to admit its shortcomings but very often, a good checklist is only the start to many a great thing. As the authors themselves admitted: “While we have tried our very best to ensure that the contents are accurate, the scale of this work means that mistakes and omissions are inevitable. We encourage everyone to verify the names in the Systema Brachyurorum whenever possible, preferably with the original literature, and to let us know of any mistakes, errors, and other problems they may encounter. This compilation represents a first attempt, and we hope to continue improving and updating it as we can.”. To accompany this publication, a “Crabs of the World” (COTW) website was created to link the public to that Supplement as well as the changes and corrections that have needed to be added since it was published, in the form of corrigenda. The improved versions of papers from the Bulletin dealt with in the article will soon be available via this portal, which we hope will make it a ‘one stop shop’ for anyone needing this literature. CHANGES TO DATES OF PUBLICATION In recent research done by the authors and E. Dickinson (Low et al., in press), several volumes of the Bulletin of Raffles Museum and the Bulletin of the National Museum have had their actual dates of publication changed. The date on these volumes does not correspond with the actual dates they were released. This is based on a close examination of the relevant wrappers of several sets of the Bulletin at various locations. A total of 13 volumes are affected by these changes, but only four are relevant to the brachyuran crab research (Volumes 9, 21, 23 and 25). A total of 17 species, described between 1935 and 1951 are affected. the discoveries only affect the available dates of the affected taxa by a year. 292 In the LITERATURE CITED, we present a complete list of all papers published in the Bulletin related to the Brachyura. ACKNOWLEDGMENTS The authors would like to acknowledge the assistance of Edward Dickinson for independently collaborating the dates of publication of the Raffles Museum and National Museum Bulletins in the United Kingdom, as well as his comments on this manuscript. Thanks are also due the librarians that have given us access to old copies of the Raffles Museum and National Museum Bulletins in their care and for helping with the checking of dates of publication. THE RAFFLES BULLETIN OF ZOOLOGY 2009 LITERATURE CITED Ahyong, S. T. & P. K. L. Ng, 2007a. Pleistacantha ori, a new species of deep-water spider crab (Crustacea: Decapoda: Brachyura: Majidae) from the Western Indian Ocean. Raffles Bulletin of Zoology, Supplement No. 16: 67–74, Figs. 1–6. Ahyong, S. T. & P. K. L. Ng, 2007b. The pinnotherid type material of Semper (1880), Nauck (1880) and Bürger (1895) (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, Supplement No. 16: 191–226, Figs. 1–26, Tables 1, 2. Ahyong, S. T. & P. K. L. Ng, 2007c. Visayeres acron, a new genus and species of pinnotherid crab (Crustacea: Decapoda: Brachyura) from the Philippines. Raffles Bulletin of Zoology, Supplement No. 16: 187–189, Fig. 1. Ahyong, S. T. & P. K. L. Ng, 2009. The Cymonomidae of the Philippines (Crustacea: Decapoda: Brachyura), with descriptions of four new species. Raffles Bulletin of Zoology, Supplement No. 20: 233–246, Figs. 1–7. Bahir, M. M. & D. C. J. Yeo, 2005. A revision of the genus Oziothephusa Müller, 1887 (Crustacea: Decapoda: Parathelphusidae), with descriptions of eight new species. Raffles Bulletin of Zoology, Supplement No. 12: 77–120, Figs. 1–46. Bahir, M. M. & D. C. J. Yeo, 2007. The gecarcinucid freshwater crabs of Southern India (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, Supplement No. 16: 309–354, Figs. 1–50. Bahir, M. M. & P. K. L. Ng, 2005. Descriptions of ten new species of freshwater crabs (Crustacea: Brachyura: Parathelphusidae: Ceylonthelphusa, Mahatha, Perbrinckia) from Sri Lanka. Raffles Bulletin of Zoology, Supplement No. 12: 47–75, Figs. 1–36. Chen H. L. & S. H. Fang, 1998. On a new species of Nursia (Crustacea: Decapoda: Brachyura: Leucosiidae) from Fujian Province, China. Raffles Bulletin of Zoology, 46(2): 605–608, Figs . 1, Table 1. Chen, H.-L. & P. K. L. Ng, 1999. Crabs of the Demania rotundata species group (Crustacea: Decapoda: Brachyura) from East and South China Seas, with description of a new species. Raffles Bulletin of Zoology, 47(1): 139–153, Figs. 1–8. Chen, H. L. & P. K. L. Ng, 2003. On new species of Leuscosiidae (Crustacea: Decapoda: Brachyura) from Singapore and the South China Sea. Raffles Bulletin of Zoology, 51(1): 61–69, Figs. 1–5, Table 1. Chen, H. L. & P. K. L. Ng, 2004. On two species of spider crabs of the genus Doclea (Crustacea: Decapoda: Brachyura: Majidae: Pisinae) from China, one of which is new. Raffles Bulletin of Zoology, 52(1): 201–208, Figs. 1–5. Chen, H. L. & Z. X. Xu, 1992. Harrovia ngi, a replacement name for Harrovia longipes Chen & Xu, 1991, preoccupied by Harrovia albolineata longipes Lanchester, 1900 (Crustacea: Decapoda: Brachyura: Eumedonidae). Raffles Bulletin of Zoology, 40(2): 265–266. Chen, W.-J., J.-H. Cheng & J.-Y. Shy, 2005. On two new species of freshwater crabs (Crustacea: Decapoda: Brachyura: Potamidae) from Southern Taiwan. Raffles Bulletin of Zoology, 53(1): 103–110, Figs. 1–6, Tables 1–4. Chia, D. G. B. & P. K. L. Ng, 1993. New records of three rare brachyuran crabs from Singapore seas (Crustacea: Decapoda: Brachyura: Parthenopidae: [sic] Xanthidae and Pilumnidae). Raffles Bulletin of Zoology, 41(1): 159–167, Fig. 1, Pl. 1, Table 1. Bahir, M. M., P. K. L. Ng, K. Crandall & R. Pethiyagoda, 2005. A conservation assessment of the freshwater crabs of Sri Lanka. Raffles Bulletin of Zoology, Supplement No. 12: 121–126, Table 1. Chia, D. G. B. & P. K. L. Ng, 1998. A revision of Ceratocarcinus White, 1847, and Harrovia Adams & White, 1849 (Crustacea: Decapoda: Eumedonidae), two genera of crabs symbiotic with crinoids. Raffles Bulletin of Zoology, 46(2): 493–563, Figs. 1–33, Tables 1, 2. Balss, H., 1938. Ueber einige Xanthidae (Crustaea Dekapoda) von Singapore und Umgebung. Bulletin of the Raffles Museum, Singapore, Straits Settlements 14: 48–63, Figs. 1–2, Pls. II, III. Chia, O. K. S. & P. K. L. Ng, 2006. The freshwater crabs of Sulawesi, with descriptions of two new genera and four new species (Crustacea: Decapoda: Brachyura: Parathelphusidae). Raffles Bulletin of Zoology, 54(2): 381–428, Figs. 1–44, Table 1. Buitendijk, A. M., 1949. On a small collection of Decapoda Brachyura, chiefly Dromiidae and Oxyrhyncha, from the neighbourhood of Singapore. Bulletin of the Raffles Museum, Singapore, 21: 59–82. Chiong, W. L. & P. K. L. Ng, 1994. The identity of Cryptopodia spatulifrons Miers, 1879, and description of a new species, Cryptopodia fistulosa (Crustacea: Decapoda: Brachyura: Parthenopidae) from Australia. Raffles Bulletin of Zoology, 42(4): 949–959, Figs. 1–5, Tables 1–2. Campos, E. & R. B. Manning, 2001. Authorship and diagnosis of the genus Arcotheres Manning, 1993 (Crustacea: Brachyura: Pinnotheridae). Raffles Bulletin of Zoology, 49(1): 167–170, Figs. 1, 2. Castro, P., 2009. Shallow-water Trapeziidae and Tetraliidae (Crustacea: Brachyura) of the Philippines (Panglao 2004 Expedition), New Guinea and Vanuatu (Santo 2006 Expedition). Raffles Bulletin of Zoology, Supplement No. 20: 271–281, Figs. 1, 2, Table 1. Castro, P., D. G. B. Chia & P. K. L. Ng, 1995. On the taxonomic status of Ceratocarcinus longimanus White, 1847 (Crustacea: Decapoda: Brachyura: Eumedonidae), a crab symbiotic with comatulid crinoids. Raffles Bulletin of Zoology, 43(1): 239–250, Fig. 1. Chen, H. L. & S. H. Fang, 1991. A new species of Leucosia (Crustacea: Brachyura: Leucosiidae) from the Taiwan Straits. Raffles Bulletin of Zoology, 39(1): 183–185, Fig. 1. Chiong, W. L. & P. K. L. Ng, 1998. A revision of the buckler crabs of the genus Cryptopodia H. Milne Edwards, 1834 (Crustacea: Decapoda: Brachyura: Parthenopidae). Raffles Bulletin of Zoology, 46(1): 157–216, Fig. 1–35, Tables 1–6. Chuang, C. T. N. & P. K. L. Ng, 1991. Preliminary descriptions of one new genus and three new species of hymenosomatid crabs from Southeast Asia (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, 39(2): 363–368, Figs. 1–2. Clark, P. F. & B. Presswell, 2001. Adam White: The crustacean years. Raffles Bulletin of Zoology, 49(1): 149–166, Tables 1, 2. Clark, P. F. & J. Paula, 2003. Descriptions of ten xanthoidean (Crustacea: Decapoda: Brachyura) first stage zoeas from Inhaca Island, Mozambique. Raffles Bulletin of Zoology, 51(2): 323–378, Figs. 1–40, Tables 1–10. Clark, P. F., P. K. L. Ng & P.-H. Ho, 2004. Atergatis subdentatus(De Haan, 1835), Atergatopsis germaini A. Milne Edwards, 1865 293 Low et al.: Brachyuran crab research in the Raffles Bulletin and Platypodia eydouxi (A. Milne Edwards, 1865) (Crustacea: Decapoda: Xanthoidea: Xanthidae: Zosiminae) – First stage zoeal descriptions with implications for the subfamily. Raffles Bulletin of Zoology, 52(2): 563–592, Figs. 1–20, Tables 1–6. Crosnier, A., 1982. Raoul Serène (1909–1980). Crustaceana, 43(2): 189–200. Crosnier, A., 2003. A new genus and species of portunid crab (Crustacea: Decapoda: Brachyura) from New Caledonia, South-West Pacific. The Raffles Bulletin of Zoology, 51(2): 315–321, Figs. 1–5. Dai, A. Y., 1997. A revision of freshwater crabs of the genus Nanhaipotamon Bott, 1968 from China (Crustacea: Decapoda: Brachyura: Potamidae). Raffles Bulletin of Zoology, 45(2): 209–235, Figs. 1–11, Pls. I, II. Dai, A.-Y. & P. K. L. Ng, 1994. Establishment of a new genus of freshwater crab, Huananpotamon (Crustacea: Decapoda: Brachyura: Potamidae) from Southern China. Raffles Bulletin of Zoology, 42(3): 657–661, Fig. 1, Table 1. Dai, A. Y. & M. Türkay, 1997. Revision of the Chinese freshwater crabs previously placed in the genus Isolapotamon Bott, 1968 (Crustacea: Decapoda: Brachyura: Potamidae). Raffles Bulletin of Zoology, 45(2): 237–264, Figs. 1–11, Pls. I, II. Dai, A. Y., X. M. Zhou & W. D. Peng, 1995. On seven new species of freshwater crabs of the genus Huananpotamon Dai & Ng, 1994 (Crustacea: Decapoda: Brachyura: Potamidae) from Jiangxi Province, Southern China. Raffles Bulletin of Zoology, 43(2): 417–433, Figs. 1–7, Pl. 1. Davie, P. J. F., 1993. Rediscovery of Camptoplax coppingeri Miers, 1884, and its placement in the Pilumnidae (Crustacea: Brachyura). Raffles Bulletin of Zoology, 41(1): 67–73, Figs. 1, 2. Davie, P. J. F., 2003. A new species of Perisesarma (Crustacea: Brachyura: Sesarmidae) from the Bay of Bengal. Raffles Bulletin of Zoology, 51(2): 387–391, Figs. 1–2. Davie, P. J. F. & N. A. Ghani, 1993. A new species of Pilumnopeus (Crustacea: Decapoda: Pilumnidae) from Pakistan. Raffles Bulletin of Zoology, 41(1): 61–65, Figs. 1, 2. Galil, B. S., 2004. A new species of Myra Leach, 1817 (Crustacea: Decapoda: Leucosiidae) from Timor Sea and South China Sea. Raffles Bulletin of Zoology, 52(1): 215–217, Figs. 1–2. Galil, B. S. & P. K. L. Ng, 2007. Leucosiid crabs from Panglao, Philippines, with description of three new species (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, Supplement No. 16: 79–94, Figs. 1–5. Galil, B. S. & P. K. L. Ng, 2009. A new species of Alox Tan & Ng, 1995 (Decapoda: Brachyura: Leucosiidae) from Balicasag Island, the Philippines. Raffles Bulletin of Zoology, Supplement No. 20: 267–270, Figs. 1–3. Gibson-Hill, C. A., 1947. Christmas Island—Field notes on terrestrial crabs. Bulletin of the Raffles Museum, 18: 43–52, Pl. III. Gordon, I., 1938. On three species of Portunidae (Decapoda, Brachyura) from the Malay Peninsula. Bulletin of the Raffles Museum, 14: 175–185, Figs. 1–6. Guinot, D., D. Doumenc, C. C. Chintiroglou, 1995. A review of the carrying behaviour in brachyuran crabs, with additional information on the symbioses with sea anemones. Raffles Bulletin of Zoology, 43(2): 377–416, Figs. 1–8, Pls. 1–3. Guo, J. Y., N. K. Ng, A. Dai & P. K. L. Ng, 1997. The taxonomy of three commercially important species of mitten crabs of the genus Eriocheir De Haan, 1835 (Crustacea: Decapoda: Brachyura: Grapsidae). Raffles Bulletin of Zoology, 45(2): 445–476, Figs. 1–9, Tables 1, 2. Harminto, S. & P. K. L. Ng, 1991. A revision of the camptandriine genus Baruna Stebbing, 1904 (Crustacea: Brachyura: Decapoda: Ocypodidae), with descriptions of two new species from the Indo-West Pacific. Raffles Bulletin of Zoology, 39(1): 187–207, Figs. 1–8. Ho, P.-H., H.-P. Yu & P. K. L. Ng, 2000. New records of Eriphiidae, Pilumnidae and Xanthidae (Crustacea: Decapoda: Brachyura) from Taiwan. Raffles Bulletin of Zoology, 48(1): 111–122, Fig. 1, Table 1. Davie, P. J. F. & T. Kosuge, 1995. A new species of Tmethypocoelis (Crustacea: Brachyura: Ocypodidae) from Japan. Raffles Bulletin of Zoology, 43(1): 207–215, Figs. 1–4, Table 1. Hsueh. P.-W. & P. K. L. Ng, 2008. Two unrecorded genera and species of brachyuran crabs (Decapoda: Camptandriidae, Pilumnidae) from Taiwan, with a note on Cancer semicylindricus Fabricius, 1798. Raffles Bulletin of Zoology, Supplement No. 19: 99–103, Figs. 1–4. Davie, P. J. F. & N. K. Ng, 2007a. Two new subfamilies of Varunidae (Crustacea: Brachyura), with description of two new genera. Raffles Bulletin of Zoology, Supplement No. 16: 257–272, Figs. 1–9. Keenan, C. P., P. J. F. Davie & D. L. Mann, 1998. A revision of the genus Scylla De Haan, 1833 (Crustacea: Decapoda: Brachyura: Portunidae). Raffles Bulletin of Zoology, 46(1): 217–245, Figs. 1–14, Tables 1–5. Davie, P. J. F. & P. K. L. Ng, 2007b. A new genus for the cavedwelling crabs previously assigned to Sesarmoides (Crustacea: Decapoda: Brachyura: Sesarmidae). Raffles Bulletin of Zoology, Supplement No. 16: 227–231, Figs. 1, 2. Kitaura, J. & K. Wada, 2006. New species of Ilyoplax (Brachyura: Ocypodidae: Dotillinae) from the Philippines and Indonesia: Behavioral, molecular, and morphological evidence. Raffles Bulletin of Zoology, 54(2): 373–379, Figs. 1–6. [31 August 2006] Davie, P. J. F. & V. X. Nguyen, 2003. A new species of Metaplax (Crustacea: Brachyura: Varunidae: Cyclograpsinae) from Vietnam. Raffles Bulletin of Zoology, 51(2): 379–386, Figs. 1–3. Davie, P. J. F. & P. A. Turner, 1994. A new record and new species of Parthenope from Northern Australia (Crustacea: Brachyura: Parthenopidae). Raffles Bulletin of Zoology, 42(4): 975–981, Figs. 1–4. Freitag, H. & D. C. J. Yeo, 2004. Two new species of Parathelphusa H. Milne Edwards, 1853, from the Philippines (Crustacea: Decapoda: Brachyura: Parathelphusidae). Raffles Bulletin of Zoology, 52(1): 227–237, Figs. 1–8, Tables 1A, 1B. 294 Koh, S. K. & P. K. L. Ng, 2000. A revision of the spiny crabs of the genus Hypothalassia Gistel, 1848 (Crustacea: Decapoda: Brachyura: Eriphiidae). Raffles Bulletin of Zoology, 48(1): 123–141, Figs. 1–12. Koh, S. K. & P. K. L. Ng, 2008. A revision of the shore crabs of the genus Eriphia (Crustacea: Brachyura: Eriphiidae). Raffles Bulletin of Zoology, 56(2): 327–355, Figs. 1–19. Komai, T. & K. Wada, 2008. A revision of the estuarune crab genus Ilyograpsus Barnard, 1955 (Crustacea: Decapoda: Brachyura: Macrophthalmidae), with descriptions of a new genus and one new species. Raffles Bulletion of Zoology, 56(2): 357–384, Figs. 1–18, Table 1. THE RAFFLES BULLETIN OF ZOOLOGY 2009 Kropp, R. K., 1994. The gall crabs (Crustacea: Decapoda: Brachyura: Cryptochiridae) of the Rumphius Expeditions revisited, with descriptions of three new species. Raffles Bulletin of Zoology, 42(3): 521–538, Figs. 1–5. Naiyanetr, P., 1994a. On three new genera of Thai ricefield crabs allied to Somannithelphusa Bott, 1968 (Crustacea: Decapoda: Brachyura: Parathelphusidae). Raffles Bulletin of Zoology, 42(3): 695–700, Figs. 1–5. Lai, J. C. Y. & B. S. Galil, 2007. A new species of Matuta Weber, 1795 (Crustacea, Decapoda, Matutidae) from Java, Indonesia. Raffles Bulletin of Zoology, Supplement No. 16: 75–77, Figs. 1–2. Naiyanetr, P., 1994b. On two species of terrestrial crabs of the genus Dromothelphusa Naiyanetr, 1992 (Crustacea: Decapoda: Brachyura: Potamidae) from Thailand. Raffles Bulletin of Zoology, 42(3): 689–694, Figs. 1, 2, Table 1. Lim, S. S. L., P. S. Lee & C. H. Diong, 2005. Influence of biotope characteristics on the distribution of Uca annulipes (H. Milne Edwards, 1837) and U. vocans (Linnaeus, 1758) (Crustacea: Brachyura: Ocypodidae) on Pulau Hantu Besar, Singapore. Raffles Bulletin of Zoology, 53(1): 111–114, Figs. 1, 2, Tables 1, 2. Naiyanetr, P. & A.-Y. Dai, 1997. On eleven new species of freshwater crabs of the genus Somanniathelphusa Bott, 1968, from Southern China (Crustacea: Decapoda: Brachyura: Parathelphusidae). Raffles Bulletin of Zoology, 45(1): 73–96, Figs. 1–11, Pls. I, II. Litulo, C., 2005. External factors determining the reproductive periodicity in a tropical population of the hairy crab Pilumnus vespertilio (Decapoda: Brachyura: Pilumnidae). Raffles Bulletin of Zoology, 53(1): 115–118, Figs. 1, 2, Tables 1, 2. Loh, L. W. & P. K. L. Ng, 1999a. A revision of the spider crabs of the genus Paratymolus Miers, 1879, with descriptions of two new genera and six new species (Crustacea: Decapoda: Brachyura: Majidae). Raffles Bulletin of Zoology, 47(2): 365–407, Figs. 1–15. Loh, L. W. & P. K. L. Ng, 1999b. Notes of some Southeast Asian species of spider crabs of the genera Doclea and Hyastenus(Crustacea: Decapoda: Brachyura: Majidae). Raffles Bulletin of Zoology, 47(1): 59–72, Figs. 1–8. Loh, L. W. & S.-H. Wu, 1998. On a new species of Paratymolus from Taiwan (Crustacea: Decapoda: Brachyura: Majidae). Raffles Bulletin of Zoology, 46(2): 609–614, Figs. 1, 2. Low, M. E. Y., S. H. Tan & P. K. L. Ng, 2009. Date of publication of the Bulletin of the Raffl es Museum and Bulletin of the National Museum, predecessors to the Raffles Bulletin of Zoology. Raffles Bulletin of Zoology (in press). Manning, R. B., 1991. The status of Cleistostoma dotilliforme Alcock, 1900 (Crustacea: Decapoda: Ocypdidae), with the description of a new genus. Raffles Bulletin of Zoology, 39(2): 299–306, Figs. 1–4. Manning, R. B., 1993. A new deep-sea crab, genus Chaceon, from Indonesia (Crustacea: Decapoda: Geryonidae). Raffles Bulletin of Zoology, 41(2): 169–172, Figs. 1, 2. Manning, R. B. & K. J. Reed, 2006. Decapod crustaceans deposited in the Zoological Museum of Copenhagen by William Stimpson in 1859. Raffles Bulletin of Zoology, 54(2): 283–293. Manuel-Santos, M. R. & P. K. L. Ng, 2007. On the genus ˇ ˇ 2005, from the Philippines and Japan, Ladomedaeus Števcic, ˇ ˇ 2005 (Decapoda: and the status of the Ladomedaeidae Števcic, Brachyura: Xanthoidea). Raffles Bulletin of Zoology, Supplement No. 16: 177–185, Figs. 1–5. [31 December 2007] McLay, C. L. & P. K. L. Ng, 2007. Revision of the Indo-West Pacific sponge crabs of the genus Petalomera Stimpson, 1858 (Decapoda: Brachyura: Dromiidae). Raffles Bulletin of Zoology, 55(1): 107–120, Figs. 1–9, Tables 1, 2. Mendoza, J. C. E. & P. K. L. Ng, 2008. New genera and species of euxanthine crabs (Crustacea: Decapoda: Brachyura: Xanthidae) from the Bohol Sea, the Philipphines. Raffles Bulletin of Zoology, 56(2): 385–404, Figs. 1–9. Naiyanetr, P., 1992. Validation of five new species of Thai potamid crabs (Crustacea: Decapoda: Brachyura). The Raffles Bulletin of Zoology, 40(1): 1–7, Fig. 1. Naruse, T. & Z. Jaafar, 2009. Geosesarma aedituens, a new terrestrial crab (Crustacea: Decapoda: Brachyura: Sesarmidae) from Bali, Indonesia. Raffles Bulletin of Zoology, 57(1): 183–187, Figs. 1–3. Naruse, T. & P. K. L. Ng, 2007a. On a new species of cavernicolous crab of the genus Sesarmoides Seréne & Soh, 1970 (Crustacea: Decapoda: Brachyura: Sesarmidae) from Sulawesi, Indonesia. Raffles Bulletin of Zoology, 55(1): 127–130, Figs. 1, 2, Table 1. Naruse, T. & P. K. L. Ng, 2007b. On a new species of Elamenopsis from Singapore, with notes on Crustaenia palawanensis (Serène, 1971) (Crustacea: Decapoda: Brachyura: Hymenosomatidae). Raffles Bulletin of Zoology, 55(1): 121–125, Figs. 1, 2. Naruse, T. & P. K. L. Ng, 2007c. Scandarma splendidum, a new species of tree-climbing crab (Decapoda: Brachyura: Sesarmidae) from Sarawak, Malaysia, Borneo. Raffles Bulletin of Zoology, 55(2): 337–341, Figs. 1–3. Naruse, T., S. Shokita & J.-Y. Shy, 2004. A new species of the freshwater crab, previously assigned to Geothelphusa miyazakii (Miyake & Chiu, 1965) (Crustacea: Decapoda: Potamidae), from Yaeyama Group, Southern Ryukyus, Japan. The Raffles Bulletin of Zoology, 52(2): 555–562, Fig. 1–4, Tables 1–3. Ng, N. K. & A. Y. Dai, 1997. Sinopotamon baiyanense, a new species of cavernicolous potamid crab (Crustacea: Decapoda: Brachyura: Potamidae) from Southern China. Raffles Bulletin of Zoology, 45(1): 139–144, Figs. 1, 2. Ng, N. K., P. J. F. Davie, C. D. Schubart & P. K. L. Ng, 2007. Xenograpsidae, a new family of grapsoid crabs (Crustacea: Brachyura) associated with shallow water hydrothermal vents. Raffles Bulletin of Zoology, Supplement No. 16: 233–256, Figs. 1–10, Tables 1–4. Ng, P. K. L., 1989a. Geithusa pulcher, new genus and species (Crustacea: Decapoda: Brachyura: Parathelphusidae) from Pulau Redang, Trengganu, Peninsular Malaysia. Raffles Bulletin of Zoology, 37(1 & 2): 31–36, Figs. 1, 2, Pl. 1. Ng, P. K. L., 1989b. Terrathelphusa, a new genus of semiterrestrial freshwater crabs from Borneo and Java (Crustacea: Decapoda: Brachyura: Sundathelphusidae). Raffles Bulletin of Zoology, 37(1 & 2): 116–131, Figs. 1–3, Pls. 1–4. Ng, P. K. L., 1989c. The identity of the cavernicolous freshwater crab Potamon (Thelphusa) bidiense Lanchester, 1900 (Crustacea: Decapoda: Brachyura: Gecarcinucidae) from Sarawak, Borneo, with description of a new genus. Raffles Bulletin of Zoology, 37(1 & 2): 63–72, Fig. 1, Pls. 1, 2. Ng, P. K. L., 1991. Cancrocaeca xenomorpha, new genus and species, a blind troglobitic freshwater hymenosomatid (Crustacea: Decapoda: Brachyura) from Sulawesi, Indonesia. Raffles Bulletin of Zoology, 39(1): 59–73, Figs. 1–7. 295 Low et al.: Brachyuran crab research in the Raffles Bulletin Ng, P. K. L., 1992. On a collection of freshwater crabs (Crustacea: Brachyura) from Terengganu, Peninsular Malaysia, with description of a second species of Geithusa (Parathelphusidae). Raffles Bulletin of Zoology, 40(1): 95–101, Figs. 1, 2, Table 1. Ng, P. K. L., 1993. Kraussiinae, a new subfamily for the genera Kraussia Dana, 1852, Palapedia, new genus, and Garthasia, new genus (Crustacea: Decapoda: Brachyura: Xanthidae), with descriptions of two new species from Singapore and the Philippines. Raffles Bulletin of Zoology, 41(1): 133–157, Figs. 1–8, Tables 1–3. Ng, P. K. L., 1995a. The freshwater crabs and prawns (Crustacea: Decapoda) of Bako National Park, Sarawak, Malaysia, with descriptions of one new genus and three new species. The Raffles Bulletin of Zoology, 43(1): 181–205, Figs. 1–14. Ng, P. K. L., 1995b. Obituary. Michael Willmer Forbes Tweedie (1907–1993). Crustaceana, 68(1): 111–115. Ng, P. K. L., 1996. Nemoron nomas, a new genus and new species of terrestrial crab (Crustacea: Decapoda: Brachyura: Potamidae) from central Vietnam. Raffles Bulletin of Zoology, 44(1): 29–36, Figs. 1–4. Ng, P. K. L., 1997. On a new genus and four new species of freshwater crabs (Crustacea: Decapoda: Brachyura: Parathelphusidae) from Borneo and Java. Raffles Bulletin of Zoology, 45(1): 105–121, Figs. 1–12, Table 1. Ng, P. K. L., 2002a. New species of cavernicolous crabs of the genus Sesarmoides from the Western Pacific, with a key to the genus (Crustacea: Decapoda: Brachyura: Sesarmidae). Raffles Bulletin of Zoology, 50(2): 419–435, Figs. 1–17. Ng, P. K. L., 2002b. On a new species of cavernicolous Neoliomera (Crustacea: Decapoda: Brachyura: Xanthidae) from Christmas Island and Ryukyus, Japan. The Raffles Bulletin of Zoology, 50(1): 95–99, Figs. 1–5. Ng, P. K. L. & S. T. Ahyong, 2001. Brachyuran type specimens (Crustacea: Decapoda) in the MacLeay Collection, University of Sydney, Australia. Raffles Bulletin of Zoology, 49(1): 83–100, Figs. 1–6, Table 1. Ng, P. K. L. & F. Alvarez, 2000. Villalobosus, a replacement name for Stygothelphusa Alvarez and Villalobos, 1991 (Crustacea: Brachyura: Pseudothelphusidae), preoccupied by Stygothelphusa Ng, 1989 (Crustacea: Brachyura: Gecarcinucidae). Raffles Bulletin of Zoology, 48(2): 337. Ng, P. K. L., H.-L. Chen & T.-Y. Chan, 1999. Taxonomic notes on three Indo-West Pacific species of Calappa (Decapoda: Brachyura: Calappidae). Raffles Bulletin of Zoology, 47(2): 607–616, Figs. 1–3. Ng, P. K. L. & D. G. B. Chia, 1994. The genus Glyptocarcinus Takeda, 1973, with descriptions of a new subfamily, two new genera and two new species from New Caledonia (Crustacea: Decapoda: Brachyura: Xanthidae). Raffles Bulletin of Zoology, 42(3): 701–730, Figs. 1–13, Tables 1, 2. Ng, P. K. L. & D. G. B. Chia, 1997. Lophozozymus erinnyes, a new species of poisonous crabs from Australia, with notes on L. pictor (Fabricius, 1798), L. incisus (H. Milne Edwards, 1834) and L. edwardsi (Odhner, 1925) (Crustacea: Decapoda: Brachyura: Xanthidae). Raffles Bulletin of Zoology, 45(2): 419–443, Figs. 1–11, Table 1. Ng, P. K. L. & C. T. N. Chuang, 1996. The Hymenosomatidae (Crustacea: Decapoda: Brachyura) of Southeast Asia, with notes on other species. Raffles Bulletin of Zoology, Supplement No. 3: 1–82, Figs. 1–32, Tables 1, 2. 296 Ng, P. K. L. & P. F. Clark, 1994. The first stage zoea of Gaillardiellus orientalis (Odhner, 1925), with notes on the subfamily Actaeinae (Crustacea: Decapoda: Brachyura: Xanthidae). Raffles Bulletin of Zoology, 42(4): 847–857, Figs. 1–4, Table 1. Ng, P. K. L., A. Y. Dai & S. L. Yang, 1997. The Indo-West Pacific Pilumnidae X. New species and records from the South China Sea (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, 45(1): 145–159, Figs. 1–6. Ng, P. K. L. & P. J. F. Davie, 2007. On the identity of Atergatis floridus (Linnaeus, 1767) and recognition of Atergatis ocyroe (Herbst, 1901) as a valid species from the Indian Ocean (Crustacea: Brachyura: Xanthidae). Raffles Bulletin of Zoology, Supplement No. 16: 169–175, Figs. 1–4. [31 December 2007] Ng, P. K. L. & D. Guinot, 2001. On the land crabs of the genus Discoplax A. Milne Edwards, 1867 (Crustacea: Decapoda: Brachyura: Gecarcinidae), with description of a new cavernicolous species from the Philippines. The Raffles Bulletin of Zoology, 49(2): 311–338, Figs. 1–16. Ng, P. K. L., D. Guinot & P. J. F. Davie, 2008. Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Supplement No. 17: 1–286, Figs. 1–190. Ng, P. K. L., D. Guinot, & T. M. Iliffe, 1996. Revision of the anchialine varunine crabs of the genus Orcovita Ng & Tomascik, 1994 (Crustacea: Decapoda: Brachyura: Grapsidae), with descriptions of four new species. Raffles Bulletin of Zoology, 44(1): 109–134, Figs. 1–16, Table 1. Ng, P. K. L. & P.-H. Ho, 2003. On the deep-water dorippid crabs of the genus Ethusina Smith, 1884 (Crustacea: Decapoda: Brachyura) from Taiwan. Raffles Bulletin of Zoology, 51(1): 71–85, Figs. 1–15. Ng, P. K. L. & G. S. Y. Lim, 1990. On the ecology of Harrovia albolineata Adams & White, 1848 (Crustacea: Decapoda: Brachyura: Eumendonidae), a crab symbiotic with crinioids. Raffles Bulletin of Zoology, 38(2): 257–262, Table 1. Ng, P. K. L., H.-C. Liu & C. D. Schubart, 2004. Geosesarma hednon, a new species of terrestrial crab (Crustacea: Decapoda: Brachyura: Sesarmidae) from Taiwan and the Philippines. Raffles Bulletin of Zoology, 52(1): 239–249, Figs. 1–14. Ng, P. K. L. & P. Naiyanetr, 1995. Pudaengon, a new genus of terrestrial crabs (Crustacea: Decapoda: Brachyura: Potamidae) from Thailand and Laos, with descriptions of seven new species. Raffles Bulletin of Zoology, 43(2): 355–376, Figs. 1–17, Table 1. Ng, P. K. L. & T. Naruse, 2007. On two species of deep-sea homolodromoid crabs of the genus Dicranodromia (Crustacea: Decapoda: Brachyura: Homolodromiidae) from the Philippines, including one new species. Raffles Bulletin of Zoology, Supplement No. 16: 47–53, Figs. 1–8. [31 December 2007] Ng, P. K. L. & T. Naruse, 2009. On the identity of Pinnixa brevipes H. Milne Edwards, 1853, and a new species of Aphanodactylus Tesch, 1918 (Crustacea: Decapoda: Brachyura: Pinnotheroidea) from the Philippines. Raffles Bulletin of Zoology, Supplement No. 20: 283–290, Figs. 1–7, Tables, 1, 2. Ng, P. K. L. & C. D. Schubart, 2002. Haberma nanum, a new genus and new species of mangrove crab (Crustacea: Decapoda: Brachyura: Sesarmidae) from Singapore. Raffles Bulletin of Zoology, 50(2): 437–442, Figs. 1–4. Ng, P. K. L., N. Sivasothi & L. K. Wang, 1994. History, dates of publication and bibliography of the Bulletin of the Raffles Museum, Bulletin of the National Museum and the Raffles THE RAFFLES BULLETIN OF ZOOLOGY 2009 Bulletin of Zoology (1928-1993). Raffles Bulletin of Zoology, 42(1): 1-105. Ng, P. K. L. & L. W. H. Tan, 1991. Irmengardia didacta, a new freshwater crab (Crustacea: Decapoda: Brachyura: Parathelphusidae) from Johor, Peninsular Malaysia. Raffles Bulletin of Zoology, 39(1): 135–140, Figs. 1, 2, Pl. 1, Table 1. Ng, P. K. L. & T. Tomascik, 1994. Orcovita saltatrix, a new genus and species of anchialine varunine crab (Crustacea: Decapoda: Brachyura: Grapsidae) from Kakaban Island, Indonesia. Raffles Bulletin of Zoology, 42(4): 937–948, Figs. 1–7. Ng, P. K. L. & D. Wowor, 1990. Terrathelphusa adipis, new species (Crustacea: Decapoda: Brachyura: Sundathelphusidae) from Kalimantan, Borneo. Raffles Bulletin of Zoology, 38(2): 263–268, Fig. 1, Pl. 1. Ng, P. K. L. & C. M. Yang, 1989. On some species of Demania Laurie, 1906 (Crustacea: Decapoda: Brachyura: Xanthidae) from Malaysia, Singapore and the Philippines, with a key for the genus. Raffles Bulletin of Zoology, 37(1 & 2): 37–50, Figs. 1, 2, Pls. 1, 2. Shen, C.-J., 1937. Notes on a collection of swimming crabs (Portunidae) from Singapore. Bulletin of the Raffles Museum, 13: 96–139, Figs. 1–20. Shokita, S., T. Naruse & H. Fuji, 2002. Geothelphusa miyakoensis, a new species of freshwater crab (Crustacea: Decapoda: Brachyura: Potamidae) from Miyako Island, Southern Ryukyus, Japan. Raffles Bulletin of Zoology, 50(2): 443–448, Figs. 1–3. Shy, J.-Y., 2005. A new species of freshwater crab of the genus Geothelphusa Stimpson, 1858 (Crustacea, Decapoda, Brachyura, Potamidae) from Taiwan. Raffles Bulletin of Zoology, 53(1): 99–102, Figs. 1–5. Shy, J.-Y., W.-J. Chen & J.-H. Cheng, 2000. Redescription of the freshwater crab Geothelphusa neipu Chen, Cheng & Shy, 1998 (Crustacea: Decapoda: Brachyura: Potamidae) from Southern Taiwan. Raffles Bulletin of Zoology, 48(1): 147–151, Figs. 1, 2, Tables 1–4. Ng, P. K. L. & C. M. Yang, 1989. On Michael Willmer Forbes Tweedie. Raffles Bulletin of Zoology, 37(1 & 2): 160-167. Shy, J.-Y., P. K. L. Ng & H.-P. Yu, 1994. Crabs of the genus Geothelphusa Stimpson, 1858 (Crustacea: Decapoda: Brachyura: Potamidae) from Taiwan, with descriptions of 25 new species. Raffles Bulletin of Zoology, 42(4): 781–846, Figs. 1–33, Tables 1, 2. Richer de Forges, B. & P. K. L. Ng, 2007a. New records and new species of Homolidae De Haan, 1839, from the Philippines and French Polynesia (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, Supplement No. 16: 29–45, Figs. 1–9. Siddiqui, F. A., Q. A. Kazmi & N. M. Tirmizi, 2000. Developmental stages of the spider crab Schizophrys pakistanensis Tirmizi & Kazmi, 1995 (Crustacea: Decapoda: Majidae). Raffles Bulletin of Zoology, 48(1): 17–29, Figs. 1–8, Tables 1–3. Richer de Forges, B. & P. K. L. Ng, 2007b. Notes on deep-sea spider crabs of the genus Cyrtomaia Miers, 1886, from the Philippines (Crustacea: Decapoda: Brachyura: Majidae), with description of a new species. Raffles Bulletin of Zoology, Supplement No. 16: 55–65, Figs. 1–6, List 1. Smedley, N, 1931. A fresh-water crab, Parathelphusa sexpunctatum(Lanch.) in the Malay Peninsula. Bulletin of the Raffles Museum, 5: 68–70, Figs. 1, 2, Pl. III. Richer de Forges, B. & P. K. L. Ng, 2009. On the majoid genera Oxypleurodon Miers, 1886, and Sphenocarcinus A MilneEdwards, 1875 (Crustacea: Brachyura: Epialtidae), with descriptions of two new genera and five new species. Raffles Bulletin of Zoology, Supplement No. 20: 247–266, Figs. 1–12, Table 1. Roux, J., 1935. New freshwater decapod crustaceans from the Malay Peninsula. Bulletin of the Raffles Museum, 9: 28–33, Figs. 1, 2, Pl. IV. Roux, J., 1936. Second note upon freshwater decapod Crustaceans from the Malay Peninsula. Bulletin of the Raffles Museum 12: 29–43, Figs. 1–14, Pls. XII, XIII. Schubart, C. D., J. I. González-Gordillo, N. B. Reyns. H.-C. Liu & J. A. Cuesta, 2001. Are Atlantic and Indo-Pacific populations of the rafting crab, Plagusia depressa (Fabricius), distinct? New evidence from larval morphology and mtDNA. Raffles Bulletin of Zoology, 49(2): 301–310, Figs. 1–5, Tables 1–4. Schubart, C. D., H.-C. Liu & J. A. Cuesta, 2003. A new genus and species of tree-climbing crab (Crustacea: Brachyura: Sesarmidae) from Taiwan with notes on its ecology and larval morphology. Raffles Bulletin of Zoology, 51(1): 49–59, Figs. 1–8. Schubart, C. D. & P. K. L. Ng, 2000. On the identities of the rafting crabs Cancer depressus Fabricius, 1775, Cancer squamosus Herbst, 1790, Plagusia immaculata Lamarck, 1818, and Plagusia tuberculata Lamarck, 1818 (Crustacea: Decapoda: Brachyura: Plagusiidae). Raffles Bulletin of Zoology, 48(2): 327–336, Figs. 1–4, Table 1. Serene, R. & C. L. Soh, 1967. A new species of Sesarma from Singapore. Bulletin of the National Museum Singapore, 33(16): 107–110, Fig. 1, Table 1. Tan C. G. S. & J.-F. Huang, 1995. Baruna sinensis, a new camptandriine crab from Taiwan (Crustacea: Decapoda: Brachyura: Ocypodidae). Raffles Bulletin of Zoology, 43(2): 447–452, Figs. 1–3. Tan, C. G. S. & A. Humpherys, 1995. Paracleistostoma tweediei, a new species of camptandriine crab, from Singapore and Brunei mangroves (Crustacea: Decapoda: Brachyura: Ocypodidae). Raffles Bulletin of Zoology, 43(1): 251–256, Figs. 1–3. Tan, C. G. S. & P. K. L. Ng, 1999. A revision of the genus Camptandrium Stimpson, 1858 (Crustacea: Decapoda: Brachyura: Camptandriidae). Raffles Bulletin of Zoology, 47(1): 193–219, Figs. 1–6, Table 1. Tan, C. G. S. & B. Richer de Forges, 1993. On the systematics and ecology of two species of mimetic crabs belonging to the family Leucosiidae (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, 41(1): 119–132, Figs. 1–6. Tan, S. H. & P. K. L. Ng, 2007a. Descriptions of new genera from the subfamily Parthenopidae (Crustacea: Decapoda: Brachyura: Parthenopidae). Raffles Bulletin of Zoology, Supplement No. 16: 95–119, Figs. 1–17. Tan, S. H. & P. K. L. Ng, 2007b. Review of the subfamily Daldorfiinae Ng & Rodríguez, 1986 (Crustacea: Decapoda: Brachyura: Parthenopidae). Raffles Bulletin of Zoology, Supplement No. 16: 121–167, Figs. 1–30. Türkay, M. & A. Y. Dai, 1997. Review of the Chinese freshwater crabs previously placed in the genus Malayopotamon Bott, 1968 (Crustacea: Decapoda: Brachyura: Potamidae). Raffles Bulletin of Zoology, 45(2): 189–207, Figs. 1–7, Pl. I. Tweedie, M. W. F., 1935. Notes on the genus Ilyoplax Stimpson (Brachyura, Ocypodidae). Bulletin of the Raffles Museum 10: 53–61, Pls. II, III. 297 Low et al.: Brachyuran crab research in the Raffles Bulletin Tweedie, M. W. F., 1936a. Note on Parathelphusa (Liothelphusa) kadamaiana. Bulletin of the Raffles Museum, 12: 70–72, Fig. 1. Wagner, H. P., 1992. A new species of Austrolibinia from the South China Sea and Indonesia (Crustacea: Brachyura: Majidae). Raffles Bulletin of Zoology, 40(1): 69–73, Figs. 1–7. Tweedie, M. W. F., 1936b) On the crabs of the family Grapsidae in the collection of the Raffles Museum. Bulletin of the Raffles Museum, 12: 44–70, Figs. 1–3, Pls. XIV, XV. Ward, M., 1935. Notes on a collection of crabs from Christmas Island, Indian Ocean. Bulletin of the Raffles Museum, 9: 5–28, Pls. I–III. Tweedie, M. W. F., 1937. On the crabs of the family Ocypodidae in the collection of the Raffles Museum. Bulletin of the Raffles Museum, 13: 140–170, Figs. 1–9. Wee, D. P. C. & P. K. L. Ng, 1995. Swimming crabs of the genera Charybdis De Haan, 1833, and Thalamita Latreille, 1829 (Crustacea: Decapoda: Brachyura: Portunidae) from Peninsular Malaysia and Singapore. Raffles Bulletin of Zoology, Supplement No. 1: 1–128. Tweedie, M. W. F., 1938a. A new scopimerine crab from the Malay Peninsula. Bulletin of the Raffles Museum, 14: 198–202, Figs. 1, 2. Tweedie, M. W. F., 1938b. Note on Parathelphusa maculata de Man. Bulletin of the Raffles Museum, 14: 203–205. Tweedie, M. W. F., 1940. New and interesting Malaysian species of Sesarma and Utica (Crustacea, Brachyura). Bulletin of the Raffles Museum, 16: 88–113, Figs. 1–12, Pl. XXIV. Tweedie, M. W. F., 1949. A collection of crabs from Aor Islands, South China Sea. Bulletin of the Raffles Museum, Singapore, 21: 83–96, Figs. 1–3. Yang, S.-L., A.-Y. Dai & P. K. L. Ng, 1998. Globopilumnus fragaria, a new species of eriphiid crab (Crustacea: Decapoda: Brachyura: Xanthoidea) from the South China Sea, with a key to the genus Globopilumnus Balss, 1933. Raffles Bulletin of Zoology, 46(1): 93–99, Figs. 1, 2. Yeo, D. C. J., 2004. A new species of Esanthelphusa (Crustacea: Brachyura: Parathelphusidae) from Laos, and a redescription of Potamon (Parathelphusa) dugasti Rathbun, 1902. Raffles Bulletin of Zoology, 52(1): 219–226, Figs. 1–5. Tweedie, M. W. F., 1950. The fauna of the Cocos-Keeling Islands, Brachyura and Stomatopoda. Bulletin of the Raffles Museum, 22: 105–148, Figs. 1–4, Pls. 16, 17. Yeo, D. C. J., Y. Cai & P. K. L. Ng, 1999. The freshwater and terrestrial decapod crustacea of Pulau Tioman, Peninsular Malaysia. Raffles Bulletin of Zoology, Supplement No. 6: 197–244, Figs. 1–20. Tweedie, M. W. F., 1951a. Notes on grapsoid crabs from the Raffles Museum. 1. A new genus and description of a new species of the subfamily Sesarminae. Bulletin of the Raffles Museum, 23: 310–316, Fig. 1. Yeo, D. C. J. & P. K. L. Ng, 1998. Freshwater crabs of the Potamon tannanti species group (Crustacea: Decapoda: Brachyura: Potamidae) from Northern Indochina. Raffles Bulletin of Zoology, 46(2): 627–650, Figs. 1–7, Tables 1, 2. Tweedie, M. W. F., 1951b. Notes on grapsoid crabs from the Raffles Museum. 2. On the habits of three Ocypodid crabs. Bulletin of the Raffles Museum, 23: 317–324, Pl. 7. Yeo, D. C. J. & P. K. L. Ng, 1999. A new species of Geosesarma (Crustacea: Decapoda: Brachyura: Grapsidae) from Pulau Tioman, Peninsular Malaysia. Raffles Bulletin of Zoology, Supplement No. 6: 189–196, Figs. 1–3. Tweedie, M. W. F., 1955a. Notes on Grapsoid Crabs from the Raffles Museum. 3. Faunal Differentiation in the regions east and west of the Malay Peninsula. Bulletin of the Raffles Museum, 25: 118–121, Fig. 1. Tweedie, M. W. F., 1955b. Notes on Grapsoid Crabs from the Raffles Museum. 4. Auditory and visual signalling. Bulletin of the Raffles Museum, 25: 121–127, Fig. 2. Tweedie, M. W. F., 1955c. Notes on Grapsoid Crabs from the Raffles Museum. 5. Metopograpsus gracilipes de Man. Bulletin of the Raffles Museum, 25: 127–128. 298 Yeo, D. C. J. & P. K. L. Ng, 2007. On the genus “Potamon” and allies in Indochina (Crustacea: Decapoda: Brachyura: Potamidae). Raffles Bulletin of Zoology, Supplement No. 16: 273–308, Figs. 1–19. Yeo, D. C. J., D. W. Rahayu & P. K. L. Ng, 2004. Brachyura (Crustacea) of the Anambas Expedition 2002. Raffles Bulletin of Zoology, Supplement No. 11: 79–88, Figs. 1, 2. THE RAFFLES BULLETIN OF ZOOLOGY 2009 Table 1. Brachyuran crab species published in the Raffles Bulletin of Zoology (and the Bulletin of the Raffles Museum and the Bulletin of the National Museum). Highlighted species indicate that the year of publication of that species has changed. Only the revised dates are shown. No. Species Remarks 1 Atergatis? tweediei Ward, 1935 2 Chlorodopsis natalensis Ward, 1935† 3 Danielea noelensis (Ward, 1935)[Medaeus] 4 Echinoecus sculptus (Ward, 1935) 5 Etisus albus (Ward, 1935) [Etisodes] 6 Hyastenus macrospinosus Ward, 1935† 7 Ilyoplax obliqua Tweedie, 1935 8 Ilyoplax punctata Tweedie, 1935 9 Kraussia proporcellana Ward, 1935† 10 Leptodius planus Ward, 1935 11 Pachygrapsus natalensis Ward, 1935† = Pachygrapsus plicatus (H. Milne Edwards, 1837) 12 Paraxanthias haematostictus Ward, 1935† = Lachnopodus ponapensis (Rathbun, 1907) 13 Percnon demani Ward, 1935 Replacement name; = Percnon planissimum (Herbst, 1804) 14 Pseudoliomera natalensis Ward, 1935† = Pseudoliomera granosimana (A. Milne-Edwards, 1865) 15 Stoliczia chaseni (Roux, 1935)[Potamon (Potamiscus)] = Liocarpilodes harmsi (Balss, 1934) [Proechinoecus] = Tylocarcinus dumerilii (H. Milne Edwards, 1834) = Kraussia rugulosa (Krauss, 1843) 16 Stoliczia tweediei (Roux, 1935)[Potamon (Potamiscus)] 17 Tweediea noelensis Ward, 1935*† = Tweediea odhneri (Gordon, 1934) 18 Episesarma singaporense (Tweedie, 1936b) [Sesarma] 19 Irmengardia pilosimana (Roux, 1936) [Paratelphusa (Peritelphusa)] 20 Johora johorensis (Roux, 1936)[Potamon (Potamon)] 21 Neosesarma gemmiferum (Tweedie, 1936b) [Sesarma] 22 Pachygrapsus quadratus Tweedie, 1936b† = Metopograpsus quadridentatus Stimpson, 1858 23 Parasesarma rutilimanum (Tweedie, 1936b) [Sesarma] 24 Perisesarma lanchesteri (Tweedie, 1936b) [Sesarma] 25 Stoliczia cognata (Roux, 1936)[Potamon (Potamiscus)] 26 Stoliczia pahangensis (Roux, 1936) 27 Stoliczia rafflesi (Roux, 1936)[Potamon (Potamiscus)] 28 Ilyogynnis microcheirum (Tweedie, 1937) 29 Ilyoplax longicarpa Tweedie, 1937 [Potamon (Potamiscus)] [Paracleistostoma] 30 Macrophthalmus (Macrophthalmus) malaccensis Tweedie, 1937 31 Macrophthalmus malayensis Tweedie, 1937† 32 Paracleistostoma longimanum Tweedie, 1937 33 Portunus (Lupocycloporus) minutus (Shen, 1937) [Neptunus (Lupocycloporus)] 34 Portunus (Xiphonectes) tweediei (Shen, 1937) [Neptunus (Hellenus)] 35 Pseudogelasimus plectodactylus Tweedie, 1937* 36 Uca (Tubuca) rosea (Tweedie, 1937) 37 Atergatopsis tweediei Balss, 1938 38 Portunus (Achelous) octodentatus (Gordon, 1938) 39 Potamocypoda pugil Tweedie, 1938* 40 Thalamita malaccensis Gordon, 1938 41 Zalasius sakai Balss, 1938 42 Bresedium sedilense (Tweedie, 1940) [Sesarma] 43 Episesarma versicolor (Tweedie, 1940) [Sesarma] 44 Geosesarma penangense (Tweedie, 1940) = Macrophthalmus (Macrophthalmus) laevimanus H. Milne Edwards, 1852 [Gelasimus] [Neptunus] [Sesarma] † 45 Perisesarma indiarum (Tweedie, 1940) 46 Pseudosesarma (“”) johorense (Tweedie, 1940) 47 Neoliomera ovata Tweedie, 1949 Replacement name; [Sesarma (Perisesarma)] [Sesarma] 299 Low et al.: Brachyuran crab research in the Raffles Bulletin Table 1. Continued. No. 48 Species Remarks Lachnopodus gibsonhilli (Tweedie, 1950) [Paraxanthias] † 49 Platypodia keelingi Tweedie, 1950 = Platypodia granulosa (Rüppell, 1830) 50 Sesarma sigallatum Tweedie, 1950† = Parasesarma carolinense Rathbun, 1907 51 Nanosesarma nunongi Tweedie, 1951* 52 Episesarma chentongense (Serène & Soh, 1967) 53 Geithusa pulchra Ng, 1989a* 54 Coccusa adipis (Ng & Wowor, 1990) [Terrathelphusa] 55 Baruna mangromurphia Harminto & Ng, 1991† = Baruna trigranulum (Dai & Song, 1986) 56 Baruna minuta Harminto & Ng, 1991 57 Cancrocaeca xenomorpha Ng, 1991* 58 Elamena globosa Chuang & Ng, 1991 59 Elamena mendosa Chuang & Ng, 1991 60 Irmengardia didacta Ng & L. W. H. Tan, 1991 61 Leucosia longimaculata Chen & Fang, 1991 62 Limnopilos naiyanetri Chuang & Ng, 1991* 63 Austrolibinia pincerna Wagner, 1992 64 Doimon maehongsonense (Naiyanetr, 1992) [Potamon] 65 Eosamon phuphanense (Naiyanetr, 1992) [Potamon] 66 Geithusa lentiginosa Ng, 1992 67 Harrovia ngi Chen & Xu, 1992†Replacement name 68 Stelomon erawanense (Naiyanetr, 1992) [Potamon] 69 Stelomon kanchanaburiense (Naiyanetr, 1992) [Potamon] 70 Takpotamon maesotense (Naiyanetr, 1992) [Potamon] 71 Chaceon karubar Manning, 1993 72 Oreotlos etor C. G. S. Tan & Richer de Forges, 1993 73 Palapedia serenei Ng, 1993* 74 Palapedia valentini Ng, 1993* 75 Pilumnopeus pereiodontus Davie & Ghani, 1993 76 Antrocarcinus petrosus Ng & Chia, 1994 77 Cryptopodia fistulosa Chiong & Ng, 1994 78 Fizesereneia latisella Kropp, 1994 79 Fizesereneia tholia Kropp, 1994 80 Geothelphusa albogilva Shy, Ng & Yu, 1994 81 Geothelphusa ancylophallus Shy, Ng & Yu, 1994 82 Geothelphusa bicolor Shy, Ng & Yu, 1994 83 Geothelphusa caesia Shy, Ng & Yu, 1994 84 Geothelphusa cinerea Shy, Ng & Yu, 1994 85 Geothelphusa dolichopodes Shy, Ng & Yu, 1994 86 Geothelphusa eucrinodonta Shy, Ng & Yu, 1994 87 Geothelphusa eurysoma Shy, Ng & Yu, 1994 88 Geothelphusa ferruginea Shy, Ng & Yu, 1994 89 Geothelphusa gracilipes Shy, Ng & Yu, 1994 90 Geothelphusa ilan Shy, Ng & Yu, 1994 91 Geothelphusa lanyu Shy, Ng & Yu, 1994 92 Geothelphusa lutao Shy, Ng & Yu, 1994 93 Geothelphusa monticola Shy, Ng & Yu, 1994 94 Geothelphusa nanao Shy, Ng & Yu, 1994 95 Geothelphusa nanhsi Shy, Ng & Yu, 1994 96 Geothelphusa olea Shy, Ng & Yu, 1994 97 Geothelphusa takuan Shy, Ng & Yu, 1994 [Sesarma (Sesarma)] 300 THE RAFFLES BULLETIN OF ZOOLOGY 2009 Table 1. Continued. No. Species 98 Geothelphusa tali Shy, Ng & Yu, 1994 Remarks 99 Geothelphusa taroko Shy, Ng & Yu, 1994 100 Geothelphusa tawu Shy, Ng & Yu, 1994 101 Geothelphusa tsayae Shy, Ng & Yu, 1994 102 Geothelphusa wangi Shy, Ng & Yu, 1994 103 Geothelphusa wutai Shy, Ng & Yu, 1994 104 Geothelphusa yangmingshan Shy, Ng & Yu, 1994 105 Glyptocarcinus politus Ng & Chia, 1994 106 Lithoscaptus prionotus Kropp, 1994 107 Orcovita saltatrix Ng & Tomascik, 1994* 108 Parthenope chondrodes Davie & Turner, 1994 109 Bakousa sarawakensis Ng, 1995a* 110 Baruna sinensis C. G. S. Tan & Huang, 1995 111 Huananpotamon chongrenense Dai, Zhou & Peng, 1995 112 Huananpotamon guixiense Dai, Zhou & Peng, 1995 113 Huananpotamon lichuanense Dai, Zhou & Peng, 1995 114 Huananpotamon medium Dai, Zhou & Peng, 1995 115 Huananpotamon nanchengense Dai, Zhou & Peng, 1995 116 Huananpotamon ruijinense Dai, Zhou & Peng, 1995 117 Huananpotamon yiyangense Dai, Zhou & Peng, 1995 118 Paracleistostoma tweediei C. G. S. Tan & Humpherys, 1995† 119 Pudaengon arnamicai Ng & Naiyanetr, 1995* 120 Pudaengon hinpoon Ng & Naiyanetr, 1995* 121 Pudaengon khammouan Ng & Naiyanetr, 1995* 122 Pudaengon mukdahan Ng & Naiyanetr, 1995* 123 Pudaengon sakonnakorn Ng & Naiyanetr, 1995* 124 Pudaengon thatphanom Ng & Naiyanetr, 1995* 125 Pudaengon wanonniwat Ng & Naiyanetr, 1995* 126 Thalamita cerasma Wee & Ng, 1995 127 Thalamita spinicarpa Wee & Ng, 1995 128 Tmethypocoelis choreutes Davie & Kosuge, 1995 129 Amarinus crenulatus Ng & Chuang, 1996 130 Amarinus pumilus Ng & Chuang, 1996 131 Apechocinus streptophallus Ng & Chuang, 1996 132 Elamena magnum Ng & Chuang, 1996 133 Elamena simplidenta Ng & Chuang, 1996 134 Elamena sundaica Ng & Chuang, 1996 135 Elamenopsis comosa Ng & Chuang, 1996 136 Nemoron nomas Ng, 1996* 137 Neorhynchoplax prima Ng & Chuang, 1996 138 Orcovita angulata Ng, Guinot & Iliffe, 1996 139 Orcovita fictilia Ng, Guinot & Iliffe, 1996 140 Orcovita gracilipes Ng, Guinot & Iliffe, 1996 141 Orcovita mollitia Ng, Guinot & Iliffe, 1996 142 Bottapotamon lingchuanense Türkay & Dai, 1997* 143 Heterochelamon guangxiense Türkay & Dai, 1997* 144 Heterochelamon yangshuoense Türkay & Dai, 1997* 145 Lophozozymus erinnyes Ng & D. G. B. Chia, 1997 146 Nanhaipotamon aculatum Dai, 1997 147 Nanhaipotamon guangdongense Dai, 1997 301 = Paracleistostoma eriophorum(Nobili, 1903) Low et al.: Brachyuran crab research in the Raffles Bulletin Table 1. Continued. No. Species Remarks 148 Nanhaipotamon hepingense Dai, 1997 149 Nanhaipotamon huaanense Dai, 1997 150 Nanhaipotamon nanriense Dai, 1997 151 Nanhaipotamon pinghense Dai, 1997 152 Nanhaipotamon pingyuanense Dai, 1997 153 Nanhaipotamon wenzhouense Dai, 1997 154 Nanhaipotamon yongchuense Dai, 1997 155 Neilupotamon xinganense Dai & Türkay, 1997 156 Parathelphusa baweanensis Ng, 1997a 157 Parathelphusa quadrata Ng, 1997a 158 Pilumnus capillatus Ng, Dai & Yang, 1997 159 Pilumnus comatus Ng, Dai & Yang, 1997 160 Pilumnus rubroseta Ng, Dai & Yang, 1997 161 Pilumnus semilunaris Ng, Dai & Yang, 1997 162 Sinopotamon baiyanense N. K. Ng & Dai, 1997 163 Somanniathelphusa amoyensis Naiyanetr & Dai, 1997 164 Somanniathelphusa araeochela Naiyanetr & Dai, 1997 165 Somanniathelphusa guilinensis Naiyanetr & Dai, 1997 166 Somanniathelphusa huaanensis Naiyanetr & Dai, 1997 167 Somanniathelphusa longicaudus Naiyanetr & Dai, 1997 168 Somanniathelphusa megachela Naiyanetr & Dai, 1997 169 Somanniathelphusa nanningensis Naiyanetr & Dai, 1997 170 Somanniathelphusa yangshanensis Naiyanetr & Dai, 1997 171 Somanniathelphusa yulinensis Naiyanetr & Dai, 1997 172 Somanniathelphusa zhangpuensis Naiyanetr & Dai, 1997 173 Somanniathelphusa zhapoensis Naiyanetr & Dai, 1997 174 Terrathelphusa ovis Ng, 1997a 175 Terrathelphusa telur Ng, 1997a 176 Viaderiana sentus Ng, Dai & Yang, 1997 177 Yarepotamon breviflagellum Dai & Türkay, 1997* 178 Yarepotamon guangdongense Dai & Türkay, 1997* 179 Cryptopodia echinosa Chiong & Ng, 1998 180 Cryptopodia patula Chiong & Ng, 1998 181 Dumea taiwanicus (Loh & Wu, 1998) [Paratymolus] 182 Eupilumnus fragaria (Yang, Dai & Ng, 1998) [Globopilumnus] 183 Harrovia cognata D. G. B. Chia & Ng, 1998 184 Indochinamon cua (Yeo & Ng, 1998) [Potamon] 185 Indochinamon guttus (Yeo & Ng, 1998) [Potamon] 186 Indochinamon ou (Yeo & Ng, 1998) [Potamon] 187 Indochinamon villosum (Yeo & Ng, 1998) [Potamon] 188 Nursia hamipleopoda Chen & Fang, 1998 189 Demania unispinosa Chen & Ng, 1999 190 Geosesarma albomita Yeo & Ng, 1999 191 Litosus giraffus Loh & Ng, 1999* 192 Moguai aloutos C. G. S. Tan & Ng, 1999* 193 Paratymolus barnardi Loh & Ng, 1999 194 Paratymolus coccus Loh & Ng, 1999 195 Paratymolus cygnus Loh & Ng, 1999 196 Paratymolus griffini Loh & Ng, 1999 197 Paratymolus prolatus Loh & Ng, 1999 302 THE RAFFLES BULLETIN OF ZOOLOGY 2009 Table 1. Continued. No. Species 198 Paratymolus vannus Loh & Ng, 1999 199 Hypothalassia acerba Koh & Ng, 2000 200 Discoplax gracilipes Ng & Guinot, 2001 201 Geothelphusa miyakoensis Shokita, Naruse & Fuji, 2002 202 Haberma nanum Ng & Schubart, 2002* Remarks 203 Karstarma balicum (Ng, 2002a)[Sesarmoides] 204 Karstarma boholano (Ng, 2002a) [Sesarmoides] 205 Karstarma guamense (Ng, 2002a) [Sesarmoides] 206 Karstarma loyalty (Ng, 2002a)[Sesarmoides] 207 Karstarma sulu (Ng, 2002a)[Sesarmoides] 208 Neoliomera cerasinus Ng, 2002b 209 Ethusina alcocki Ng & Ho, 2003b† 210 Ethusina chenae Ng & Ho, 2003b 211 Ethusina insolita Ng & Ho, 2003b 212 Ethusina macrospina Ng & Ho, 2003b 213 Ethusina saltator Ng & Ho, 2003b† 214 Ethusina taiwanensis Ng & Ho, 2003b 215 Geosesarma hednon Ng, Liu & Schubart, 2003 216 Heteronucia margaritata Chen & Ng, 2003 217 Leucosia singaporensis Chen & Ng, 2003† 218 Metaplax gocongensis Davie & Nguyen, 2003 219 Perisesarma bengalense Davie, 2003 220 Pseudophilyra punctata Chen & Ng, 2003 221 Richerellus moosai Crosnier, 2003* 222 Scandarma lintou Schubart, Liu & Cuesta, 2003* 223 Soceulia major (Chen & Ng, 2003) 224 Doclea unidentata Chen & Ng, 2004 225 Esanthelphusa nimoafi Yeo, 2004 226 Geothelphusa fulva Naruse, Shokita & Shy, 2004 227 Geothelphusa marginata Naruse, Shokita & Shy, 2004 228 Myra digitata Galil, 2004 229 Parasesarma anambas Yeo, Rahayu & Ng, 2004 230 Parathelphusa cabayugan Freitag & Yeo, 2004 231 Parathelphusa manguao Freitag & Yeo, 2004 232 Ceylonthelphusa alpina Bahir & Ng, 2005 233 Ceylonthelphusa diva Bahir & Ng, 2005 234 Ceylonthelphusa durrelli Bahir & Ng, 2005 235 Ceylonthelphusa savitriae Bahir & Ng, 2005 236 Geothelphusa leeae Shy, 2005 237 Geothelphusa lili Chen, Cheng & Shy, 2005 238 Geothelphusa shernshan Chen, Cheng & Shy, 2005 239 Mahatha helaya Bahir & Ng, 2005 240 Mahatha lacuna Bahir & Ng, 2005 241 Mahatha regina Bahir & Ng, 2005 242 Oziotelphusa biloba Bahir & Yeo, 2005 243 Oziotelphusa dakuna Bahir & Yeo, 2005 244 Oziotelphusa gallicola Bahir & Yeo, 2005 245 Oziotelphusa intuta Bahir & Yeo, 2005 246 Oziotelphusa kerala Bahir & Yeo, 2005 247 Oziotelphusa kodagoda Bahir & Yeo, 2005 = Ethusina robusta (Miers, 1886) = Ethusina paralongipes Chen, 1993 = Soceulia brunnea (Miers, 1877) [Leucosia] 303 Low et al.: Brachyuran crab research in the Raffles Bulletin Table 1. Continued. No. Species Remarks 248 Oziotelphusa populosa Bahir & Yeo, 2005 249 Oziotelphusa ritigala Bahir & Yeo, 2005 250 Perbrinckia fenestra Bahir & Ng, 2005 251 Perbrinckia gabadagei Bahir & Ng, 2005 252 Perbrinckia rosae Bahir & Ng, 2005 253 Ilyoplax pacifica Kitaura & Wada, 2006 254 Migmathelphusa olivacea O. K. S. Chia & Ng, 2006* 255 Parathelphusa ferruginea O. K. S. Chia & Ng, 2006 256 Parathelphusa sorella O. K. S. Chia & Ng, 2006 257 Syntripsa flavichela O. K. S. Chia & Ng, 2006* 258 Alox bothros Galil & Ng, 2007 259 Alox chaunos Galil & Ng, 2007 260 Arcotheres rayi Ahyong & Ng, 2007b 261 Baratha peena Bahir & Yeo, 2007* 262 Baratha pushta Bahir & Yeo, 2007* 263 Cyrtomaia largoi Richer de Forges & Ng, 2007a 264 Daldorfia calconopia Tan & Ng, 2007 265 Daldorfia dimorpha Tan & Ng, 2007 266 Dicranodromia chenae Ng & Naruse, 2007 267 Elamenopsis rotunda Naruse & Ng, 2007b 268 Homola poupini Richer de Forges & Ng, 2007b 269 Karstarma microphthalmus (Naruse & Ng, 2007a) 270 Ladomedaeus fungillus Manuel-Santos & Ng, 2007 271 Latreillopsis mariveneae Richer de Forges & Ng, 2007b 272 Matuta purnama Lai & Galil, 2007 273 Pilarta anuka Bahir & Yeo, 2007* 274 Pleistacantha ori Ahyong & Ng, 2007a 275 Scandarma splendidum Naruse & Ng, 2007c 276 Snaha aruna Bahir & Yeo, 2007* 277 Travancoriana charu Bahir & Yeo, 2007 278 Travancoriana kuleera Bahir & Yeo, 2007 279 Urnalana cristata Galil & Ng, 2007 280 Vanni ashini Bahir & Yeo, 2007* 281 Vanni deepta Bahir & Yeo, 2007* 282 Vanni giri Bahir & Yeo, 2007* 283 Vela virupa Bahir & Yeo, 2007* 284 Viridotheres otto Ahyong & Ng, 2007b 285 Visayeres acron Ahyong & Ng, 2007c* 286 Yaldwynopsis guinotae Richer de Forges & Ng, 2007b 287 Yaldwynopsis saguili Richer de Forges & Ng, 2007b 288 Eriphia ferox Koh & Ng, 2008 289 Hepatoporus pumex Mendoza & Ng, 2008 290 Ilyograpsus daviei Komai & Wada, 2008 291 Rizalthus anconis Mendoza & Ng, 2008* 292 Visayax estampadori Mendoza & Ng, 2008* 293 Visayax osteodictyon Mendoza & Ng, 2008* 294 Alox tormos Galil & Ng, 2009 295 Aphanodactylus panglao Ng & Naruse, 2009 296 Cymonomus deforgesi Ahyong & Ng, 2009 297 Cymonomus diogenes Ahyong & Ng, 2009 [Sesarmoides] 304 THE RAFFLES BULLETIN OF ZOOLOGY 2009 Table 1. Continued. No. Species Remarks 298 Cymonomus gracilipes Ahyong & Ng, 2009 299 Cymonomus mariveneae Ahyong & Ng, 2009 300 Geosesarma aedituens Naruse & Jaafar, 2009 301 Oxypleurodon alaini Richer de Forges & Ng, 2009 302 Oxypleurodon barazeri Richer de Forges & Ng, 2009 303 Oxypleurodon boholense Richer de Forges & Ng, 2009 304 Oxypleurodon parallelum Richer de Forges & Ng, 2009 305 Stegopleurodon planirostrum Richer de Forges & Ng, 2009* Notes: 1. An * after the species denotes that the genus was erected for the species described. 2. A † after the species denotes that it is now a junior synonym. 3. Genera in brackets “[]” denote the original genus. 4. An = sign preceding a species denotes the senior synonym of the species. Table 2. Published valid species described in Crustaceana, the Journal of Crustacean Biology and the Raffles Bulletin (and its predecessors). First year of publication Age (years) Descriptions of new species Synonymised names Valid descriptions Valid species descriptions per year Crustaceana 1960 50 95 7 88 1.76 Journal of Crustacean Biology 1981 29 60 7 53 1.83 RBZ 1928 82 302* 17 285 3.48 Zootaxa 2001 10 59 0 59 5.90 Journal *Excludes three replacement names from the total of 305 names published. Table 3. Brachyuran crab genera published in the Raffles Bulletin of Zoology, and the Bulletin of the Raffles Museum and the Bulletin of the National Museum. Genera with grey background indicate that the year of publication of that genera has changed. Only the revised dates are shown. No. Genus 1 Actiomera Ng, Guinot & Davie, 2008 2 Agolambrus Tan & Ng, 2007a 3 Alcomon Yeo & Ng, 2007 4 Antrocarcinus Ng & D. G. B. Chia, 1994 5 Apograpsus Komai & Wada, 2008 6 Aspermon Yeo & Ng, 2007 7 Badistemon Yeo & Ng, 2007 8 Bakousa Ng, 1995a 9 Baratha Bahir & Yeo, 2007 10 Beccumon Yeo & Ng, 2007 11 Bottapotamon Turkay & Dai, 1997 12 Cancrocaeca Ng, 1991 13 Celatopesia Chiong & Ng, 1998 14 Chulathelphusa Naiyanetr, 1994a* 15 Costalambrus Tan & Ng, 2007a 16 Crustaenia Ng & Chuang, 1996 17 Cyrtocarcinus Ng & D. G. B. Chia, 1994 18 Derilambrus Tan & Ng, 2007a 19 Distolambrus Tan & Ng, 2007a Remarks Replacement name. = Mekhongthelphusa Naiyanetr, 1985 305 Low et al.: Brachyuran crab research in the Raffles Bulletin Table 3. Continued. No. Genus 20 Doimon Yeo & Ng, 2007 21 Dolos Tan & Richer de Forges, 1993 22 Dumea Loh & Ng, 1999a 23 Eosamon Yeo & Ng, 2007 24 Esanthelphusa Naiyanetr, 1994a 25 Garthasia Ng, 1993 26 Geithusa Ng, 1989a 27 Haberma Ng & Schubart, 2002 28 Heikeopsis Ng, Guinot & Davie, 2008 29 Heterochelamon Turkay & Dai, 1997 30 Huananpotamon Dai & Ng, 1994 31 Hypolambrus Tan & Ng, 2007a 32 Indochinamon Yeo & Ng, 2007 33 Inlethelphusa Yeo & Ng, 2007 34 Iomon Yeo & Ng, 2007 35 Karstama Davie & Ng, 2007b 36 Kempamon Yeo & Ng, 2007 37 Kukrimon Yeo & Ng, 2007 38 Lamella Bahir & Yeo, 2007 39 Latopotamon Dai & Turkay, 1997 40 Latulambrus Tan & Ng, 2007a 41 Limnopilos Chuang & Ng, 1991 42 Litosus Loh & Ng, 1999a 43 Mediapotamon Turkay & Dai, 1997 44 Megacephalomon Yeo & Ng, 2007 45 Migmathelphusa O. K. S. Chia & Ng, 2006 46 Minpotamon Dai & Turkay, 1997 47 Moguai Tan & Ng, 1999 48 Nanosesarma Tweedie, 1951a 49 Nanusia Tan & Ng, 1999 50 Nasima Manning, 1991 51 Neilupotamon Dai & Turkay, 1997 52 Nemoron Ng, 1996 53 Niobafia Tan & Ng, 2007a 54 Nodolambrus Tan & Ng, 2007a 55 Ochtholambrus Tan & Ng, 2007a 56 Olenorfia Tan & Ng, 2007a 57 Orcovita Ng & Tomascik, 1994 58 Palapedia Ng, 1993 59 Patulambrus Tan & Ng, 2007a 60 Permanotus D. G. B. Chia & Ng, 1998 61 Pilarta Bahir & Yeo, 2007 62 Piloslambrus Tan & Ng, 2007a 63 Planumon Yeo & Ng, 2007 64 Potamocypoda Tweedie, 1938a 65 Proechinoecus Ward, 1935* 66 Pseudogaetice Davie & N. K. Ng, 2007 67 Pseudogelasimus Tweedie, 1937 68 Pudaengon Ng & Naiyanetr, 1995 69 Pupamon Yeo & Ng, 2007 Remarks Replacement name. = Echinoecus Rathbun, 1894 306 THE RAFFLES BULLETIN OF ZOOLOGY 2009 Table 3. Continued. No. 70 Genus Remarks Quadramon Yeo & Ng, 2007 71 Rhinocarcinus Richer de Forges & Ng, 2009 72 Richerellus Crosnier, 2003 73 Rizalthus Mendoza & Ng, 2008 74 Sayamia Naiyanetr, 1994a 75 Scandarma Schubart, Liu & Cuesta, 2003 76 Sestrostoma Davie & N. K. Ng, 2007 77 Setosamon Yeo & Ng, 2007 78 Shanphusa Yeo & Ng, 2007 79 Snaha Bahir & Yeo, 2007 80 Spinolambrus Tan & Ng, 2007a 81 Stegopleurodon Richer de Forges & Ng, 2009 82 Stygothelphusa Ng, 1989c 83 Syntripsa O. K. S. Chia & Ng, 2006 84 Takedellus Tan & Ng, 1999 85 Tauropus D. G. B. Chia & Ng, 1998 86 Teretamon Yeo & Ng, 2007 87 Terrathelphusa Ng, 1989b 88 Thalassograpsus Tweedie, 1950 89 Tiaramedon D. G. B. Chia & Ng, 1998 90 Torhusa Ng, 1997 91 Tweediea Ward, 1935 92 Vadosapotamon Dai & Turkay, 1997 93 Vanni Bahir & Yeo, 2007 94 Vela Bahir & Yeo, 2007 95 Velolambrus Tan & Ng, 2007a 96 Villalobosus Ng & Alvarez, 2000 97 Visayax Mendoza & Ng, 2008 98 Visayeres Ahyong & Ng, 2007c 99 Yarepotamon Dai & Turkay, 1997 Replacement name Notes: 1. An * after the genus denotes that the genus has been synonymised. 2. An = sign preceding a species denotes the senior synonym of the species. 307 View publication stats